E-Flora BC: Electronic Atlas of the Flora of British Columbia

Summary:
Features include 1) a large, convex to flat or depressed, leathery fruitbody with whitish to brownish colors tinted with pink, reddish, or orange, somewhat zoned and often cracked, the margin incurved and lobed, 2) flesh that is white to light brown and not zoned, 3) decurrent, crowded teeth, 4) a stem that is central to off-center, bent or irregularly shaped, often bifurcate or trifurcate, and colored like the cap, 5) single to concrescent growth under conifers, and 6) microscopic characters. |Both the Baird(1) and the Baird(2) descriptions are based on dried material. |Some species in Sarcodon were reassigned starting in 2019 to Hydnellum based largely on molecular criteria (Larsson(2)); Larsson(2) introduced spore length (with ornamentation excluded) as a possible differentiating criterion between the two genera: 7.4-9 microns for Sarcodon and 4.45-6.95 microns for Hydnellum, with Sarcodon quercinofibulatus as an exception. If this criterion holds, Sarcodon stereosarcinon is likely to be moved to Hydnellum.

Sarcodon stereosarcinon is found in NS and NC. A collection from BC determined by K.A. Harrison is deposited at Pacific Forestry Centre.

Cap:
up to 18.5cm across, flat to depressed, irregular, lobed, leathery; pale orange, flesh, red-hair color, to camel, somalis to fawn at the disc; subzonate; pubescent to bald, often cracked or the surface burrowed by insects exposing the inner flesh, (Baird(1)), up to 8cm across, convex to flat or depressed; white, Sahara or darker to dark brown; "tomentose, radially rugose, smooth to areolate exposing inner flesh"; margin incurved, lobed; pileoli [small accessory caps] rare, (Baird(2)), 2-14(18)cm across, very irregular, light brown darkening to dull brown, faintly zonate, in humid weather sometimes exuding a light yellow juice, (Smith), 2-24(18)cm across, "single or fused, very irregular, convex", flat, rarely funnel-shaped, margin irregular, undulating [wavy], drooping, thin, incurved; light brown darkening to dull brown, faintly zoned, "occasionally in humid weather exuding light yellow juice", margin brownish on bruising; surface "smooth or finely scaly, sometimes cracked in dry weather", (Harrison), pale orange-white when young, yellowish golden brown, light brown to dark brown when old; bald to finely pubescent, surface often diffusely scattered with tiny warts, (Franklin)

Flesh:
up to 0.5cm thick, "not duplex, azonate"; white to light brown; in stem "not duplex, zonate, concolorous with pileus flesh", (Baird(1)), "context not duplex, azonate", white; in stem, "not duplex, azonate", white or tinted with brown, (Baird(2)), brownish white (Smith), firm, fibrous, scissile; brownish white; for stem flesh see STEM, (Harrison)

Teeth:
up to 0.4cm long, decurrent to base of stem, crowded, pale orange to red-hair color or somalis, often brown with lighter colored tips, very few pale blue, (Baird(1)), up to 0.4cm long, highly decurrent, often spathulate, colored as the stem or lighter toward the margin, pale blue areas rare, (Baird(2)), up to 1cm long (Smith), up to 1cm, decurrent, fine to very coarse, subulate [awl-shaped], or spathulate near stem, concolorous, tips lighter and concolorous with the growing margin, (Harrison)

Stem:
up to 6cm x 1.2cm, central to eccentric [off-center], round in cross-section to narrowing in lower part to a rooting base, bent or irregularly shaped, often bifurcate or trifurcate; colored as cap; scabrous from abortive spines to bald, (Baird(1)), up to 4.5cm x 1.2cm, central to eccentric or lateral, round in cross-section, narrowing in lower part to a point; colored as cap; tomentose toward base to matted or bald in upper part, scabrous from aborted spines, (Baird(2)), 4-8cm x 1-2cm, "tapering to a point, base of young specimens at times with droplets of reddish brown juice", (Smith), 4-8cm x 1-2cm, "irregular, tapering to a point, concolorous, flesh firm and fibrous, light brown, sometimes grayish (lilac in Coker and Beers, 1951) toward base of stipe; base of young plants occasionally showing droplets of reddish-brown juice", (Harrison)

Chemical Reactions:
context tissue olivaceous to nonreactive in KOH, (Baird(1)), preserved context olive green in KOH (Baird(2))

Odor:
none (Baird(1), Harrison), absent to weakly pungent or farinaceous, (Franklin)

Taste:
none (Baird(1)), slightly farinaceous (Smith), slight, farinaceous, (Harrison), absent to mildly pungent (Franklin)

Microscopic:
spores 4.5-5.3 x 3.5-4.5 microns, nearly round to oblong, brownish; ornamentation tuberculate, angular nodulose, hilar appendage oblique; basidia 4-spored, 22.5-32.0 x 5.0-6.0 microns, clavate, without clamp connections, sterigmata 3.0-4.8 microns long; spine trama hyphae up to 6.5(11.0) microns wide, uninflated to inflated, without clamp connections; cap trama hyphae up to 20 microns wide, inflated, interwoven throughout the context, without clamp connections, gloeoplerous-like hyphae up to 9.0 microns wide; stem hyphae up to 12.5 microns wide, inflated, interwoven throughout the context, without clamp connections, gloeoplerous-like hyphae up to 7.5 microns wide, skeletalized generative hyphae present, (Baird(1)), spores 4.5-5.3 x (3.6)4.0-4.9 microns, nearly round to oblong, "angular, nodulose, brownish", "ornamentation tuberculate", hilar appendage oblique; basidia 4-spored, 22.5-31.0 x 4.0-5.6 microns, clavate, without clamp connection, sterigmata 3.0-4.1 microns long; cap trama hyphae up to 8.5 microns wide, interwoven throughout context, inflated to uninflated, without clamp connections, gloeoplerous-like hyphae present; stem hyphae up to 8.0 microns wide, interwoven throughout the context, inflated to uninflated, without clamp connections, gloeoplerous-like hyphae absent, skeletalized generative hyphae present, (Baird(2)), spores 4.5-5 x 3.5-4.5 microns (Smith), 4.5-5.0 x 3.5-4.5 microns, nearly round to oblong, tuberculate, (Harrison)

Spore Deposit:
[presumably brown]